Rafiq, K., Jordan, N. R., Meloro, C., Wilson, A. M., Hayward, M. W., Wich, S. A., & McNutt, J. W. Scent-marking strategies of a solitary carnivore: boundary and road scent marking in the leopard. Animal Behaviour, 161, 115-126. https://doi.org/10.1016/j.anbehav.2019.12.016